Home Publications

63. Brabec, J., Rochat, E., Knudsen, R., Scholz, T. & Blasco-Costa, I. Mining various genomic resources to resolve old alpha-taxonomy questions: A test of the species hypothesis of the Proteocephalus longicollis species complex (Cestoda: Platyhelminthes) from salmonid fishes. International Journal for Parasitology 53, 197–205. PDF

62. Llopis-Belenguer, C., Balbuena, J. A., Blasco-Costa, I., Karvonen, A., Sarabeev, V. &  Jokela, J. 2023. Sensitivity of bipartite network analyses to incomplete sampling and taxonomic uncertainty. Ecology 104, e3974. PDF


61. Rochat, E., Paterson, R., Blasco-Costa, I., Power, M., Adams, C. E., Greer, R. & Knudsen, R. 2022. Temporal stability of polymorphic Arctic charr parasite communities reflect sustained divergent trophic niches. Ecology and Evolution 12, e9460. PDF

60. Feijen, F., Zajac, N., Vorburger, C., Blasco-Costa, I. & Jokela, J. 2022. Phylogeography and cryptic species structure of a locally adapted parasite in New Zealand. Molecular Ecology 31, 4112–4126. PDF


59. Blasco-Costa, I., Hayward, A., Poulin, R., & Balbuena, J. A. 2021. Next-generation cophylogeny: unravelling eco-evolutionary processes. Trends in Ecology & Evolution 36, 907-918.PDF

58. Rochat, E. C., Brodersen, J., & Blasco-Costa, I. 2021. Conspecific migration and environmental setting determine parasite infracommunities of non-migratory individual fish. Parasitology 148, 1057-1066. PDF

57. Balbuena, J. A., Monlleó-Borrull, C., Llopis-Belenguer, C., Blasco-Costa, I., Sarabeev, V. L., & Morand, S. 2021. Fuzzy quantification of common and rare species in ecological communities (FuzzyQ). Methods in Ecology and Evolution 12, 1070-1079. Link


56. Llopis-Belenguer, C., Pavoine, S., Blasco-Costa, I. & Balbuena, J.A. 2020. Assembly rules of helminth parasite communities in grey mullets: combining components of diversity. International Journal for Parasitology 50, 1089-1098. Link

55. Balbuena, J.A., Pérez-Escobar, O.A., Llopis-Belenguer, C. & Blasco-Costa, I. 2020. Random Tanglegram Partitions (Random TaPas): An Alexandrian approach to the cophylogenetic Gordian knot. Systematic Biology 69, 1212–1230. Link

54. Rochat, E.C., Blasco-Costa, I., Scholz, T. & Unmack, P.J. 2020. High diversity of metazoan parasites in carp gudgeons (Eleotridae: Hypseleotris spp.) from Eastern Australia. Journal of Helminthology 94, e146. Link

53. Llopis-Belenguer, C., Blasco-Costa, I., Balbuena, J.A., Sarabeev, V. & Stouffer, D.B. 2020. Native and invasive hosts play different roles in host-parasite networks. Ecography 43, 559–568. PDF

52. Presswell, B. & Blasco-Costa, I. 2020. Description of Tylodelphys darbyi n. sp. (Trematoda: Diplostomidae) from the threatened Australasian crested grebe (Podiceps cristatus australis, Gould 1844) and linking of its life-cycle stages. Journal of Helminthology 94, e40. Link

51. Blasco-Costa, I., Seppälä, K., Feijen, F., Zajac, N., Klappert, K. & Jokela,J. 2020. A new species of Atriophallophorus Deblock & Rosé, 1964 (Trematoda: Microphallidae) described from in vitro-grown adults and metacercariae from Potamopyrgus antipodarum (Gray, 1843) (Mollusca: Tateidae). Journal of Helminthology 94, e108. Link


50. Llopis-Belenguer, C., Balbuena, J.A., Lange, K., de Bello, F. & Blasco-Costa, I. 2019. Towards a unified functional trait framework for parasites. Trends in Parasitology 35, 972–982. Link

49. Radačovská, A., Bazsalovicsová, E., Blasco-Costa, I., Orosová, M., Gustinelli, A. & Králová-Hromadová, I. 2019. Occurrence of Dibothriocephalus latus in European perch from Alpine lakes, an important focus of diphyllobothriosis in Europe. Revue suisse de Zoologie 126, 219–225. PDF

48. Villar-Torres, M., Repullés-Albelda, A., Montero, F.E., Raga, J.A. & Blasco-Costa, I. 2019. Neither Diplectanum nor specific: A dramatic twist to the taxonomic framework of Diplectanum (Monogenea: Dactylogyridae). International Journal for Parasitology 49, 365–374. Link

47. Mendoza-Palmero, C.A., Blasco-Costa, I. & Pérez-Ponce de León, G. 2019. Morphological and molecular characterisation of a new species of Gyrodactylus von Nordmann, 1832 (Monogenoidea: Gyrodactylidae) of cichlid fishes (Perciformes) from Mexico. Parasitology International 70, 102–111. Link

46. Paterson, R.A., Knudsen, R., Blasco-Costa, I., Dunn, A.M., Hytterød, S. & Hansen, H. 2019. Determinants of parasite distribution in Arctic charr populations: catchment structure versus dispersal potential. Journal of Helminthology 93, 559 – 566. Link


45. Llopis-Belenguer, C., Blasco-Costa, I. & Balbuena, J.A. 2018. Beyond counting species: A new way to look at biodiversity. Mètode Science Studies Journal 98, 47–51. PDF

44. Llopis-Belenguer, C., Blasco-Costa, I. & Balbuena, J.A. 2018. Evaluation of three methods for biomass estimation in small invertebrates, using three large disparate parasite species as model organisms. Scientific Reports 8, 3897. PDF

43. Padros, F., Knudsen, R. & Blasco-Costa, I. 2018. Histopathological characterisation of retinal lesions associated to Diplostomum species (Platyhelminthes: Trematoda) infection in polymorphic Arctic charr Salvelinus alpinus. International Journal for Parasitology-Parasites and Wildlife 7, 68-74. PDF

42. Galaktionov, K.V. & Blasco-Costa, I. 2018 Microphallus ochotensis sp. nov.(Digenea, Microphallidae) and relative merits of two-host microphallid life cycles. Parasitology Research 117, 1051-1068. PDF

41. Acosta, A.A., Scholz, T., Blasco-Costa, I., Alves, P.V. & da Silva, R.J. 2018. A new genus and two new species of dactylogyrid monogeneans from gills of Neotropical catfishes (Siluriformes: Doradidae and Loricariidae). Parasitology International 67, 4-12. Link


40. Blasco-Costa, I. & Locke, S. A. 2017. Life history, systematics and evolution of the Diplostomoidea Poirier, 1886: progress, promises and challenges emerging from molecular studies. Advances in Parasitology, 98,167–225.

39. Blasco-Costa, I. & Poulin, R. 2017. Parasite life-cycle studies: a plea to resurrect an old parasitological tradition. Journal of Helminthology, 91, 647-656.doi: 10.1017/S0022149X16000924

38. Blasco-Costa, I., Poulin, R. & Presswell, B. 2017. Morphological description and molecular analyses of Tylodelphys sp. (Trematoda: Diplostomidae) newly recorded from the freshwater fish Gobiomorphus cotidianus (common bully) in New Zealand. Journal of Helminthology, 91, 332-345.doi: 10.1017/S0022149X16000298

37. Georgieva, S., Blasco-Costa, I. & Kostadinova, A. 2017. Molecular characterisation of four echinostomes (Digenea: Echinostomatidae) from birds in New Zealand, with descriptions of Echinostoma novaezealandense n. sp. and Echinoparyphium poulini n. sp. Systematic Parasitology, 94, 477-497.

36. Mendoza-Palmero, C.A., Blasco-Costa, I., Hernández-Mena, D. & Pérez-Ponce de León, G. 2017. Parasciadicleithrum octofasciatumn. gen., n. sp. (Monogenoidea: Dactylogyridae), parasite of Rocio octofasciata (Regan) (Cichlidae: Perciformes) from Mexico characterised by morphological and molecular evidence. Parasitology International,66, 152-162.


35. Niebuhr, C.N. & Blasco-Costa, I. 2016. Improving detection of avian malaria from host blood: a step towards a standardised protocol for diagnostics. Parasitology Research, 115, 3905-3911.

34. Lagrue, C., Joannes, A., Poulin, R. & Blasco-Costa, I. 2016. Genetic structure and host–parasite co-divergence: evidence for trait-specific local adaptation. Biological Journal of the Linnean Society, 118, 344-358.

33. Poulin, R., Blasco-Costa, I. & Randhawa, H. S. 2016. Integrating parasitology and marine ecology: Seven challenges towards greater synergy. Journal of Sea Research,113, 3-10.

32. Blasco-Costa, I., Poulin, R. & Presswell, B. 2016. Species of Apatemon Szidat, 1928 and Australapatemon Sudarikov, 1959 (Trematoda: Strigeidae) from New Zealand: linking and characterising life cycle stages with morphology and molecules. Parasitology Research,115, 271-289.

31. Blasco-Costa, I., Cutmore, S.C., Miller, T.L. & Nolan, M.J. 2016. Molecular approaches to trematode systematics: ‘best practice’ and implications for future study. Systematic Parasitology, 93, 295-306.

30. Hofmann, H., Blasco-Costa, I., Knudsen, R., Matthaei, C.D., Valois, A. & Lange, K. 2016. Parasite prevalence in an intermediate snail host is subject to multiple anthropogenic stressors in a New Zealand river system. Ecological Indicators, 60, 845-852.


29. Mendoza-Palmero, C.A., Blasco-Costa, I. & Scholz, T. 2015. Molecular phylogeny of Neotropical monogeneans (Platyhelminthes: Monogenea) from catfishes (Siluriformes). Parasites & Vectors, 8, 164.

28. Blasco-Costa, I., Rouco, C. & Poulin, R. 2015. Biogeography of parasitism in freshwater fish: spatial patterns in hot spots of infection. Ecography, 38, 301-310.


27. Faltynkova, A., Georgieva, S., Kostadinova, A., Blasco-Costa, I., Scholz, T. & Skirnisson, K. 2014. Diplostomum von Nordmann, 1832 (Digenea: Diplostomidae) in the sub-Arctic: descriptions of the larval stages of six species discovered recently in Iceland. Systematic Parasitology, 89, 195-213.

26. Herrmann, K.K., Poulin, R., Keeney, D.B. & Blasco-Costa, I. 2014. Genetic structure in a progenetic trematode: signs of cryptic species with contrasting reproductive strategies. International Journal for Parasitology, 44, 811-818.

25. Blasco-Costa, I.,Georgieva, S., Faltýnková, A., Skirnisson, K., Scholz, T. & Kostadinova, A. 2014. Fish pathogens in the Subarctic: a molecular glimpse of the diversity of Diplostomum. International Journal of Parasitology, 44, 703-715.

24. O’Dwyer, K., Blasco-Costa, I., Poulin, R. & Faltýnková, A. 2014. Four marine digenean parasites of Austrolittorina spp. (Gastropoda: Littorinidae) in New Zealand: morphological and molecular data. Systematic Parasitology, 89, 133-152.

23. Presswell, B., Blasco-Costa, I. & Kostadinova, A. 2014. Two new species of Maritrema Nicoll, 1907 (Digenea: Microphallidae) from New Zealand: morphological and molecular characterisation. Parasitology Research, 113, 1641-1656.

22. Georgieva, S., Faltynkova, A., Brown, R., Blasco-Costa, I., Soldanova, M., Sitko, J., Scholz, T. & Kostadinova, A. 2014. Echinostomarevolutum‘ (Digenea: Echinostomatidae) species complex revisited: species delimitation based on novel molecular and morphological data gathered in Europe. Parasites & Vectors, 7, 520.


21. Blasco-Costa, I. & Poulin, R. 2013. Host traits explain the genetic structure of parasites: a meta-analysis. Parasitology, 140, 1316-1322.

20. Balbuena, J.A., Míguez-Lozano, R. & Blasco-Costa, I. 2013. PACo: a novel Procrustes application to cophylogenetic analysis. PLoS ONE, 8 (4), e61048.

19. Hernández-Orts, J. S., Aznar, F. J., Blasco-Costa, I., García, N. A., Víllora-Montero, M., Crespo, E. A., Raga, J. A. & Montero, F. E. 2013. Description, microhabitat selection and infection patterns of sealworm larvae (Pseudoterranova decipiens species complex, Nematoda: Ascaridoidea) in fishes from Patagonia, Argentina. Parasites & Vectors, 6, 252.

18. Chibwana, F.D., Blasco-Costa, I., Georgieva, S., Hosea, K.M., Nkwengulila, G., Scholz, T. & Kostadinova, A. 2013. A first insight into the barcodes for African diplostomiasis: Cryptic species complex in Clarias gariepinus (Siluriformes: Clariidae). Infection, Genetics and Evolution, 17, 62-70.

17. Blasco-Costa, I., Koehler, A.V., Martin, A. & Poulin, R. 2013. Upstream-downstream gradient in infection levels by fish parasites: a common river pattern? Parasitology, 140, 266-274.


16. Blasco-Costa, I., Míguez-Lozano, R. & Balbuena, J.A. 2012. Molecular phylogeny of species of Ligophorus(Monogenea: Dactylogyridae) and their affinities within the Dactylogyridae. Parasitology International, 61, 619-627.

15. Míguez-Lozano, R., Pardo-Carranza, T.V., Blasco-Costa, I. & Balbuena, J.A. 2012. Spatial structure of helminth communities of the golden grey mullet, Liza aurata (Actinopterygii: Mugilidae), in the western Mediterranean. Journal of Parasitology, 98, 904-912.

14. Galactionov, K., Blasco-Costa, I. & Olson, P.D. 2012. Life cycles, molecular phylogeny and historical biogeography of the ‘pygmaeus’ microphallids (Digenea: Microphallidae): widespread parasites of marine and coastal birds in the Holarctic. Parasitology, 139, 1346-1360.

13. Blasco-Costa, I., Waters, J.M. & Poulin, R. 2012. Swimming against the current: genetic structure, host mobility and the drift paradox in trematode parasites. Molecular Ecology, 21, 207-217.

12. Millán, J. & Blasco-Costa, I. 2012. Molecular evidence of shared hookworm Ancylostoma tubaeformehaplotypes between the endangered Iberian lynx and sympatric domestic cats. Veterinary Parasitology, 186, 518-522.


11. Blasco-Costa, I., Balbuena, J.A., Raga, J.A., Kostadinova, A. & Olson, P.D. 2010. Molecules and morphology reveal cryptic variation among digeneans infecting sympatric mullets in the Mediterranean. Parasitology,137, 287-302. doi: 10.1017/S0031182009991375


10. Blasco-Costa, I., Balbuena, J.A, Kostadinova, A. & Olson, P.D. 2009. Interrelationships of the Haploporinae (Digenea: Haploporidae): A molecular test of the taxonomic framework based on morphology. Parasitology International, 58, 265-269. doi:10.1016/j.parint.2009.03.006

9. Blasco-Costa, I., Gibson, D.I., Balbuena, J.A., Raga, J.A. & Kostadinova, A. 2009. A revision of the Haploporinae Nicoll, 1914 (Digenea: Haploporidae) from mullets(Mugilidae): HaploporusLooss, 1902 and LecithobotrysLooss, 1902. Systematic Parasitology, 73, 107-133. doi: 10.1007/s11230-009-9175-9

8. Blasco-Costa, I., Montero, F.E., Balbuena, J.A., Raga, J.A. & Kostadinova, A. 2009. A revision of the Haploporinae Nicoll, 1914 (Digenea: Haploporidae) from mullets (Mugilidae): DicrogasterLooss, 1902 and ForticulcitaOverstreet, 1982. Systematic Parasitology, 72, 187-206. doi: 10.1007/s11230-008-9165-3

7. Blasco-Costa, I., Montero, F.E., Gibson, D.I., Balbuena, J.A. & Kostadinova, A. 2009. A revision of the Haploporinae Nicoll, 1914 (Digenea: Haploporidae) from mullets (Mugilidae): Two new haploporine genera and a key to the genera of the subfamily. Systematic Parasitology, 72, 207-215. doi: 10.1007/s11230-008-9164-4

6. Blasco-Costa, I., Montero, F.E., Gibson, D.I., Balbuena, J.A., Raga, J.A. & Kostadinova, A. 2009. A revision of the Haploporinae Nicoll, 1914 (Digenea: Haploporidae) from mullets (Mugilidae): Saccocoelium Looss, 1902. Systematic Parasitology, 72, 159-186. doi: 10.1007/s11230-008-9163-5


5. Blasco-Costa, I., Montero, F.E., Gibson, D.I., Balbuena, J.A., Raga, J.A., Shvetsova, L.S. & Kostadinova, A. 2008. A revision of SaturniusManter, 1969 (Digenea: Hemiuridae), parasites of mullets (Teleostei: Mugilidae). Systematic Parasitology,71, 53-74. doi: 10.1007/s11230-008-9141-y


4. Blasco-Costa, I., Pankov, P., Gibson, D.I., Balbuena, J.A., Raga, J.A., Sarabeev, V.L. & Kostadinova, A. 2006. Saturnius minutusn. sp. and S. dimitrovin. sp. (Digenea: Hemiuridae) from Mugil cephalusL. (Teleostei: Mugilidae), with a multivariate morphological analysis of the Mediterranean species of SaturniusManter, 1969. Systematic Parasitology, 65, 77-91. doi: 10.1007/s11230-006-9043-9

3. Holzer, A.S., Blasco-Costa, I., Sarabeev, V.L., Ovcharenko, M.O. & Balbuena, J.A. 2006. Kudoa trifoliasp. n. – Molecular phylogeny suggests a new spore morphology and unusual tissue location for a well-known genus. Journal of Fish Diseases29, 743-755. doi: 10.1111/j.1365-2761.2006.00770.x

2. Pankov, P., Webster, B.L., Blasco-Costa, I., Gibson, D.I., Littlewood, D.T.J., Balbuena, J.A. & Kostadinova, A. 2006. Robinia auratan. g., n. sp. (Digenea: Hemiuridae) from the mugilid Liza auratawith a molecular confirmation of its position within the Hemiuroidea. Parasitology, 133, 217-227. doi: 10.1017/S0031182006000126

1. Rubtsova, N., Balbuena, J.A., Sarabeev, V.L., Blasco-Costa, I. & Euzet, L. 2006. Description and morphometrical variability of a new species of Ligophorusand of Ligophoruschabaudi(Monogenea: Dactylogyridae) on Mugil cephalus(Teleostei) from the Mediterranean basin. Journal of Parasitology, 92, 486-495.  doi: 10.1645/GE-747R.1


Blasco-Costa, I. 2009. Taxonomy of the Haploporinae Nicoll, 1914 and Bunocotylinae Dollfus, 1950 (Digenea) from Mediterranean mullets (Teleostei): morphological and molecular approaches.PhD Thesis. Valencia: Universitat de Valencia. Servei de Publicacions. ISBN: 978-84-370-7564-8.